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Trees on farms are often overlooked in agricultural and natural resource research and policy in Sub-Saharan Africa. This paper addresses this gap using data from the Living Standards Measurement Study–Integrated Surveys on Agriculture in five countries: Ethiopia, Malawi, Nigeria, Tanzania, and Uganda. Trees on farms are widespread. On average, one third of rural smallholders grow trees. They account for an average of 17 percent of total annual gross income for tree-growing households and 6 percent for all rural households. Gender, land and labor endowments, and especially forest proximity and national context are key determinants of on-farm tree adoption and management. These new, national-scale insights on the prevalence, economic contribution and determinants of trees on farms in Africa lay the basis for exploring the interaction of agriculture, on-farm tree cultivation, and forestry. This will improve our understanding of rural livelihood dynamics.